Exploring Fennoscandian agricultural history through genetic analysis of aged crop materials

Exploring Fennoscandian agricultural history through genetic analysis of aged crop materials PDF Author: Maria Lundström
Publisher: Linköping University Electronic Press
ISBN: 917685194X
Category :
Languages : en
Pages : 59

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Book Description
Crop plants have undergone a multitude of genetic changes during and following their domestication. The spread of agriculture brought the crops to new geographic regions exposing them to new environments and selection pressures along the way. This gave rise to many local variants with traits favoured both by agricultural practices and the environment. Agriculture was introduced in Fennoscandia (Norway, Sweden, Finland and Denmark) around 4000 BC. The composition of the archaeobotanical record gives some clues as to which species were cultivated, but macroscale analyses rarely reach beyond that. Therefore, methods like genetic analysis are necessary to expand our knowledge about the history of crop cultivation. Under optimal conditions, DNA can survive in biological samples for several hundred thousand years. The preservation of plant specimens in the Fennoscandian climate has, however, rarely been explored. This thesis therefore attempts to dive deeper into the Fennoscandian cultivation history through genetic analyses of aged plant materials from both museum collections and archaeological sources. Cereal grains from a range of preservation conditions were evaluated to find which ones might be of interest for genetic investigations. Desiccated materials gave the highest success rates, in agreement with previous studies. Waterlogged materials appeared to contain small amounts of endogenous DNA, whereas genetic analysis of charred cereals failed completely in all samples. Population structure was investigated in 17-19th century materials of both barley and rye from Sweden and Finland. Northern and southern populations of Finnish six-row barley were distinct from one another. In southern Sweden, genetic analysis suggested conserved population structure extending over 200 years. The genetic composition of rye also seemed mostly conserved, but rye did not show geographic population structure across the investigated region in Sweden and Finland. A long-standing question in Fennoscandian crop history has been the interpretation of historical written records mentioning Brassica (cole crops, turnips and mustards), as well as the species identity of archaeobotanical finds of Brassica seeds. Thus, Next Generation Sequencing (NGS) was applied to identify which Brassica types that were cultivated in 17th century Kalmar, Sweden. The analysis corroborated morphological species classification in two of the investigated subfossil seeds, whereas no conclusions could be drawn from the remaining samples. The genome coverages were too low to allow subspecies identification. Wheat has been cultivated in Fennoscandia since the introduction of agriculture but has increased dramatically in importance over the last century. The functional allele of the wheat nutrition gene NAM-B1 was found to be particularly prominent in Fennoscandian wheats, likely associated with its effect on grain maturation time. Here the evolutionary history of NAM-B1 was investigated to see if it could truly be considered a domestication gene as suggested in a previous study. By studying extant landrace materials of Mediterranean tetraploid wheat, it was found that the non-functional allele showed signs indicative of a selective sweep. This selection did not, however, appear to have occurred during domestication. In conclusion, aged plant specimens from both museum and archaeological contexts could contribute greatly to our knowledge about historical cultivation, extending the investigated period into the mid 17th century. Subfossil and waterlogged archaeobotanical materials do contain endogenous DNA, suggesting that they are better suited for genetic analysis than charred ones, at least as far as cereals are concerned. There is potential for classifying archaeological Brassica remains using NGS, even though further optimisation of sample and library preparation may be necessary. And finally, despite NAM-B1 showing signs of selection it should not be considered a domestication gene in tetraploid wheat.

Exploring Fennoscandian agricultural history through genetic analysis of aged crop materials

Exploring Fennoscandian agricultural history through genetic analysis of aged crop materials PDF Author: Maria Lundström
Publisher: Linköping University Electronic Press
ISBN: 917685194X
Category :
Languages : en
Pages : 59

Get Book Here

Book Description
Crop plants have undergone a multitude of genetic changes during and following their domestication. The spread of agriculture brought the crops to new geographic regions exposing them to new environments and selection pressures along the way. This gave rise to many local variants with traits favoured both by agricultural practices and the environment. Agriculture was introduced in Fennoscandia (Norway, Sweden, Finland and Denmark) around 4000 BC. The composition of the archaeobotanical record gives some clues as to which species were cultivated, but macroscale analyses rarely reach beyond that. Therefore, methods like genetic analysis are necessary to expand our knowledge about the history of crop cultivation. Under optimal conditions, DNA can survive in biological samples for several hundred thousand years. The preservation of plant specimens in the Fennoscandian climate has, however, rarely been explored. This thesis therefore attempts to dive deeper into the Fennoscandian cultivation history through genetic analyses of aged plant materials from both museum collections and archaeological sources. Cereal grains from a range of preservation conditions were evaluated to find which ones might be of interest for genetic investigations. Desiccated materials gave the highest success rates, in agreement with previous studies. Waterlogged materials appeared to contain small amounts of endogenous DNA, whereas genetic analysis of charred cereals failed completely in all samples. Population structure was investigated in 17-19th century materials of both barley and rye from Sweden and Finland. Northern and southern populations of Finnish six-row barley were distinct from one another. In southern Sweden, genetic analysis suggested conserved population structure extending over 200 years. The genetic composition of rye also seemed mostly conserved, but rye did not show geographic population structure across the investigated region in Sweden and Finland. A long-standing question in Fennoscandian crop history has been the interpretation of historical written records mentioning Brassica (cole crops, turnips and mustards), as well as the species identity of archaeobotanical finds of Brassica seeds. Thus, Next Generation Sequencing (NGS) was applied to identify which Brassica types that were cultivated in 17th century Kalmar, Sweden. The analysis corroborated morphological species classification in two of the investigated subfossil seeds, whereas no conclusions could be drawn from the remaining samples. The genome coverages were too low to allow subspecies identification. Wheat has been cultivated in Fennoscandia since the introduction of agriculture but has increased dramatically in importance over the last century. The functional allele of the wheat nutrition gene NAM-B1 was found to be particularly prominent in Fennoscandian wheats, likely associated with its effect on grain maturation time. Here the evolutionary history of NAM-B1 was investigated to see if it could truly be considered a domestication gene as suggested in a previous study. By studying extant landrace materials of Mediterranean tetraploid wheat, it was found that the non-functional allele showed signs indicative of a selective sweep. This selection did not, however, appear to have occurred during domestication. In conclusion, aged plant specimens from both museum and archaeological contexts could contribute greatly to our knowledge about historical cultivation, extending the investigated period into the mid 17th century. Subfossil and waterlogged archaeobotanical materials do contain endogenous DNA, suggesting that they are better suited for genetic analysis than charred ones, at least as far as cereals are concerned. There is potential for classifying archaeological Brassica remains using NGS, even though further optimisation of sample and library preparation may be necessary. And finally, despite NAM-B1 showing signs of selection it should not be considered a domestication gene in tetraploid wheat.

Dog behaviour

Dog behaviour PDF Author: Ann-Sofie Sundman
Publisher: Linköping University Electronic Press
ISBN: 9176850722
Category :
Languages : en
Pages : 67

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Book Description
Dogs, Canis familiaris, share the lives of humans all over the world. That dogs, and the behavior of dogs, are of interest to many is therefore no surprise. In this thesis, the main aim has been to identify factors that affect dogs’ behaviours. The dog, Canis familiaris, is our first domesticated animal. Since domestication, various types of dogs have developed through adaptation to an environment shared with humans and through our selective breeding, resulting in a unique variation in morphology and behaviour. Although there is an individual variation in the behaviour of dogs, there is also a difference between breeds. Moreover, selection during the last decades has split some breeds into divergent types. Labrador and golden retrievers are divided into a common type, for show and companionship, and a field type, for hunting. By comparing the breed types, we can study the effects of recent selection. In Paper I, we investigate differences in general behavioural traits between Labrador and golden retriever and between common and field type within the two breeds by using results from the standardized behaviour test Dog Mentality Assessment. There were differences between breeds and types for all behavioural traits. However, there was also an interaction between breed and type. Thus, a common/field-type Labrador does not behave like a common/field-type golden retriever. Even though they have been selected for similar traits, the selection has affected the general behavioural traits differently in the two breeds. In paper II, we were interested in dogs’ human-directed social skills. Dogs have a high social competence when it comes to humans. Two experiments commonly used to study these skills are the problem-solving test, where dogs’ human-directed behaviours when faced with a problem are measured, and the pointing test, where dogs are tested on how well they understand human gestures. We compared the social skills of German shepherds and Labrador retrievers, and of common- and field-type Labradors. Labradors were more successful in the pointing test and German shepherds stayed closer to their owners during the problem solving. Among Labrador types, the field type had more human eye contact than the common type. Importantly, when comparing the two experiments, we found no positive correlations between the problem-solving test and the pointing test, suggesting that the two tests measure different aspects of human-directed social behaviour in dogs. A previous study has identified two suggestive genetic regions for human-directed social behaviours during the problem-solving test in beagles. In paper III, we show that these SNPs are also associated to social behaviours in Labrador and golden retrievers. Moreover, the Labrador breed types differed significantly in allele frequencies. This indicates that the two SNPs have been affected by recent selection and may have a part in the differences in sociability between common and field type. The behaviour of dogs cannot simply be explained by genetics, there is also an environmental component. In paper IV, we study which factors that affect long-term stress in dogs. Long-term cortisol can be measured by hair samples. We found a clear synchronization in hair cortisol concentrations between dogs and their owners. Neither dogs’ activity levels nor their behavioural traits affected the cortisol, however, the personality of the owners did. Therefore, we suggest that dogs mirror the stress level of their owners. The mediator between genes and the environment is epigenetics, and one epigenetic factor is DNA methylation. In paper V, we compared methylation patterns of wolves and dogs as well as dog breeds. Between both wolves and dogs and among dogs there were substantial differences in methylated DNA regions, suggesting that DNA methylation is likely to contribute to the vast variation among canines. We hypothesize that epigenetic factors have been important during domestication and in breed formation. In this thesis, I cover several aspects on how dogs’ behaviours can be affected, and paint an intricate picture on how genetics, epigenetics, and human-dog relations forms dog behaviour.

Broadening the Genetic Base of Crop Production

Broadening the Genetic Base of Crop Production PDF Author: H. David Cooper
Publisher:
ISBN:
Category : Crops
Languages : en
Pages : 484

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Book Description
This book focuses on the previously neglected interface between the conservation of plant genetic resources and their utilization. Only through utilization can the potential value of conserved genetic resources be realized. However, as this book shows, much conserved germplasm has to be subjected to long-term pre-breeding and genetic enhancement before it can be used in plant breeding programs.The authors explore the rationale and approaches for such pre-breeding efforts as the basis for broadening the genetic bases of crop production. Examples from a range of major food crops are presented and issues analyzed by leading authorities from around the world.

The Regional Impacts of Climate Change

The Regional Impacts of Climate Change PDF Author: Intergovernmental Panel on Climate Change. Working Group II.
Publisher: Cambridge University Press
ISBN: 9780521634557
Category : Science
Languages : en
Pages : 532

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Book Description
Cambridge, UK : Cambridge University Press, 1998.

The Oxford Handbook of the Archaeology and Anthropology of Hunter-Gatherers

The Oxford Handbook of the Archaeology and Anthropology of Hunter-Gatherers PDF Author: Vicki Cummings
Publisher: OUP Oxford
ISBN: 0191025275
Category : Social Science
Languages : en
Pages : 1361

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Book Description
For more than a century, the study of hunting and gathering societies has been central to the development of both archaeology and anthropology as academic disciplines, and has also generated widespread public interest and debate. The Oxford Handbook of the Archaeology and Anthropology of Hunter-Gatherers provides a comprehensive review of hunter-gatherer studies to date, including critical engagements with older debates, new theoretical perspectives, and renewed obligations for greater engagement between researchers and indigenous communities. Chapters provide in-depth archaeological, historical, and anthropological case-studies, and examine far-reaching questions about human social relations, attitudes to technology, ecology, and management of resources and the environment, as well as issues of diet, health, and gender relations - all central topics in hunter-gatherer research, but also themes that have great relevance for modern global society and its future challenges. The Handbook also provides a strategic vision for how the integration of new methods, approaches, and study regions can ensure that future research into the archaeology and anthropology of hunter-gatherers will continue to deliver penetrating insights into the factors that underlie all human diversity.

The Anthropocene as a Geological Time Unit

The Anthropocene as a Geological Time Unit PDF Author: Jan Zalasiewicz
Publisher: Cambridge University Press
ISBN: 110847523X
Category : Medical
Languages : en
Pages : 385

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Book Description
Reviews the evidence underpinning the Anthropocene as a geological epoch written by the Anthropocene Working Group investigating it. The book discusses ongoing changes to the Earth system within the context of deep geological time, allowing a comparison between the global transition taking place today with major transitions in Earth history.

Allelopathy

Allelopathy PDF Author: Manuel Joaquín Reigosa Roger
Publisher: Springer Science & Business Media
ISBN: 9781402042799
Category : Nature
Languages : en
Pages : 660

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Book Description
This book provides the reader relevant information about actual knowledge about the process of allelopathy, covering all aspects from the molecular to the ecological level. Special relevance is given to the physiological and ecophysiological aspects of allelopathy. Several ecosystems are studied and methodological considerations are taken into account in several different chapters. The book has been written to be useful both for Ph.D. students and for senior researchers, so the chapters include all necessary information to be read by beginners, but they also include a lot of useful information and discussion for the initiated.

Environment and Society in the Long Late Antiquity

Environment and Society in the Long Late Antiquity PDF Author:
Publisher: BRILL
ISBN: 9004392084
Category : History
Languages : en
Pages : 407

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Book Description
Environment and Society in the Long Late Antiquity brings together scientific, archaeological and historical evidence on the interplay of social change and environmental phenomena at the end of Antiquity and the dawn of the Middle Ages, covering the period ca. 300-800 AD. It gives a new impetus to the study of the environmental history of this crucial period of transition between two major epochs in premodern history. The volume contains both systematic overviews of the previous scholarship and available data, as well as a number of interdisciplinary case studies. It covers a wide range of topics, including the histories of landscape, climate, disease and earthquakes, all intertwined with social, cultural, economic and political developments. Contributors are Daniel Abel-Schaad , Francesca Alba-Sánchez, Flavio Anselmetti, José Antonio López-Sáez, Daniel Ariztegui, Brunhilda Brushulli, Yolanda Carrión Marco, Alexandra Chavarría, Petra Dark, Carmen Fernández Ochoa, Martin Finné, Asuunta Florenzano, Ralph Fyfe,Didier Galop, Benjamin Graham, John Haldon, Kyle Harper, Richard Hodges, Adam Izdebski, Katarina Kouli, Inga Labuhn, Tamara Lewit, Anna Maria Mercuri, Alessia Masi, Lucas McMahon, Lee Mordechai, Mario Morellón, Timothy Newfield, Almudena Orejas Saco del Valle, Leonor Peña-Chocarro, Sebastián Pérez-Díaz, Eleonora Regattieri, Stephen Rippon, Neil Roberts, Laura Sadori, Abigail Sargent, Gaia Sinopoli, Paolo Squatriti, Giovanni Stranieri, Raymond van Dam, Bernd Wagner, Mark Whittow, Penelope Wilson, Jessie Woodbridge. See inside the book.

The Oxford Handbook of the European Bronze Age

The Oxford Handbook of the European Bronze Age PDF Author: Harry Fokkens
Publisher: Oxford University Press
ISBN: 0199572860
Category : History
Languages : en
Pages : 1012

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Book Description
The Oxford Handbook of the European Bronze Age is a wide-ranging survey of a crucial period in prehistory during which many social, economic, and technological changes took place. Written by expert specialists in the field, the book provides coverage both of the themes that characterize the period, and of the specific developments that took place in the various countries of Europe. After an introduction and a discussion of chronology, successive chapters deal with settlement studies, burial analysis, hoards and hoarding, monumentality, rock art, cosmology, gender, and trade, as well as a series of articles on specific technologies and crafts (such as transport, metals, glass, salt, textiles, and weighing). The second half of the book covers each country in turn. From Ireland to Russia, Scandinavia to Sicily, every area is considered, and up to date information on important recent finds is discussed in detail. The book is the first to consider the whole of the European Bronze Age in both geographical and thematic terms, and will be the standard book on the subject for the foreseeable future.

Fundamentals of Geomorphology

Fundamentals of Geomorphology PDF Author: Richard John Huggett
Publisher: Routledge
ISBN: 1135281130
Category : Science
Languages : en
Pages : 909

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Book Description
This extensively revised, restructured, and updated edition continues to present an engaging and comprehensive introduction to the subject, exploring the world’s landforms from a broad systems perspective. It covers the basics of Earth surface forms and processes, while reflecting on the latest developments in the field. Fundamentals of Geomorphology begins with a consideration of the nature of geomorphology, process and form, history, and geomorphic systems, and moves on to discuss: structure: structural landforms associated with plate tectonics and those associated with volcanoes, impact craters, and folds, faults, and joints process and form: landforms resulting from, or influenced by, the exogenic agencies of weathering, running water, flowing ice and meltwater, ground ice and frost, the wind, and the sea; landforms developed on limestone; and landscape evolution, a discussion of ancient landforms, including palaeosurfaces, stagnant landscape features, and evolutionary aspects of landscape change. This third edition has been fully updated to include a clearer initial explanation of the nature of geomorphology, of land surface process and form, and of land-surface change over different timescales. The text has been restructured to incorporate information on geomorphic materials and processes at more suitable points in the book. Finally, historical geomorphology has been integrated throughout the text to reflect the importance of history in all aspects of geomorphology. Fundamentals of Geomorphology provides a stimulating and innovative perspective on the key topics and debates within the field of geomorphology. Written in an accessible and lively manner, it includes guides to further reading, chapter summaries, and an extensive glossary of key terms. The book is also illustrated throughout with over 200 informative diagrams and attractive photographs, all in colour.